curs pp4 sistem

7/24/2019 Curs Pp4 Sistem.

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SISTEMATICASISTEMATICA

MICROORGANISMELORMICROORGANISMELORNOIUNI DE TAXONOMIE INOIUNI DE TAXONOMIE I

NOMENCLATURNOMENCLATURBACTERIANBACTERIAN


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TAXONOMIATAXONOMIAMICROORGANISMELORMICROORGANISMELOR

TaxonomiaTaxonomia(gr. taxon = grup(gr. taxon = grup!"t! #t$$n%a!"t! #t$$n%a&'a"$&)r$$ (grup)r$$ $n%!'or* o+$!&t!'or*&'a"$&)r$$ (grup)r$$ $n%!'or* o+$!&t!'or*,!no-!n!'or*,!no-!n!'or*n a &)r!$ ",!r) /! pr!o&up)r$ $ntr)n a &)r!$ ",!r) /! pr!o&up)r$ $ntr)ur-)toar!'! o+$!&t$0!1ur-)toar!'! o+$!&t$0!1

2.2. stabilireastabilireacriteriilorcriteriilor de gruparede grupare(criterii(criterii taxonomicetaxonomice))334.4. clasifcareaclasifcareaindiviilorindiviilor"au a un$t)5$'or /!"au a un$t)5$'or /!

+a6) n+a6) n grup!grup!((taxonitaxoni p! +a6a &r$t!r$$'or p! +a6a &r$t!r$$'or"ta+$'$t!3"ta+$'$t!3

7.7. nomenclaturanomenclatura"au"au numirea taxonilornumirea taxonilor..8.8. identifcareaidentifcareaprin comparaie cu taxoniiprin comparaie cu taxoniiexisteni a unui obiect, fenomen sau in nouexisteni a unui obiect, fenomen sau in nouobservatobservat..


3/22

Cr$t!r$u' &!' -a$ o+$!&t$0 /!Cr$t!r$u' &!' -a$ o+$!&t$0 /!&'a"$&ar! a $n%!'or 0$$ !"t!&'a"$&ar! a $n%!'or 0$$ !"t!

criteriul flogenetic!criteriul flogenetic!"n"ntaxon biologictaxon biologicr!un!#t! $n/$0$6$$r!un!#t! $n/$0$6$$asem#n#tori datorit#asem#n#tori datorit#

descenden$ei dintr%un str#mo&descenden$ei dintr%un str#mo&comuncomun* po"!"or$ a$ un!$ $n,or-a%$$* po"!"or$ a$ un!$ $n,or-a%$$g!n!t$&! &o-un!.g!n!t$&! &o-un!.


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CC'a"$&ar!a'a"$&ar!a 'og!n!t$&) a'og!n!t$&) a-$&roorgan$"-!'or-$&roorgan$"-!'or a nt9-p$nat nu-!roa"!a nt9-p$nat nu-!roa"!/$&u't)5$* g!n!rat! /! &au6! o+$!&t$0!*/$&u't)5$* g!n!rat! /! &au6! o+$!&t$0!*/$ntr! &ar! -!n5$on)-1/$ntr! &ar! -!n5$on)-1:: num#rul redus de caracterenum#rul redus de caracteremor'ologicemor'ologice33:: nu pot f studiatnu pot f studiat unun indiviindividd ci ci oo

populaipopulaiee** p! &ar! o nu-$-p! &ar! o nu-$- tulpin#tulpin#clon# sau suclon# sau su!:: apariia 'recvent# *n populaiileapariia 'recvent# *n populaiilemicrobienemicrobiene a variantelora variantelorgeneticegenetice ..


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Grupar!a +a&t!r$$'or nGrupar!a +a&t!r$$'or n categorii taxonomicecategorii taxonomicepepecriterii flogeneticecriterii flogeneticea /!0!n$t po"$+$') pr$na /!0!n$t po"$+$') pr$nap'$&ar!a t!;ap'$&ar!a t!;n$&$'orn$&$'or/! +$o'og$! -o'!&u'ar) n/! +$o'og$! -o'!&u'ar) n

"tu/$$'! /! g!n!t$&) +a&t!r$an)."tu/$$'! /! g!n!t$&) +a&t!r$an).Gra/u' /! o-o'og$! a -at!r$a'u'u$ g!n!t$& a /ou)Gra/u' /! o-o'og$! a -at!r$a'u'u$ g!n!t$& a /ou)-$&roorgan$"-! (gra/u' /! nru/$r! poat! apr!&$at pr$n-$&roorgan$"-! (gra/u' /! nru/$r! poat! apr!&$at pr$n

2.2. coninutul relativ de guanin#coninutul relativ de guanin# citoin#( +citoin#( +C,) al A-NC,) al A-Npurifcat. cu c/t coninutul +purifcat. cu c/t coninutul +

C a dou#C a dou# microorganismemicroorganismeeste mai apropiat cu at/t acestea sunt mai *nrudite.este mai apropiat cu at/t acestea sunt mai *nrudite.

0!0! num#rul snum#rul secvenelor nucleotidiceecvenelor nucleotidice comune dincomune din

structurastructura A-N.A-N. 1! num#rul de secvene oligonucleotidice comune din1! num#rul de secvene oligonucleotidice comune din

ARN riboomal!ARN riboomal!

&'a"$&)r$'! &on0!n5$ona'! a'!&'a"$&)r$'! &on0!n5$ona'! a'! -$&roorgan$"-!'or +a6at!-$&roorgan$"-!'or +a6at!p!p!&r$t!r$u' "$-$'$tu/$n$$ &ara&t!r!'or&r$t!r$u' "$-$'$tu/$n$$ &ara&t!r!'orenotipiceenotipice(morologice,(morologice,culturale, biochimice, depatogenitate etc)culturale, biochimice, depatogenitate etc) au ,o"tau ,o"t"upu"! unor r!p!tat! r!-an$!r$."upu"! unor r!p!tat! r!-an$!r$.


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Categorii de taxoniCategorii de taxoni22peciapecia -- uun$tat!a taxono-$&) /! +a6) a 'u-$$n$tat!a taxono-$&) /! +a6) a 'u-$$

0$$.0$$. n &a6u' organ$"-!'or "up!r$oar!* /$nn &a6u' organ$"-!'or "up!r$oar!* /$nr!gnu' 0!g!ta' "au an$-a'* !x$"t) &r$t!r$$r!gnu' 0!g!ta' "au an$-a'* !x$"t) &r$t!r$$+$n! "ta+$'$t! /! /!'$-$tar! a "p!&$$'or* &!'+$n! "ta+$'$t! /! /!'$-$tar! a "p!&$$'or* &!'

-a$ $-portant $n/-a$ $-portant $n/ capacitatea decapacitatea de*ncruci3are*ncruci3are..22peciapecia==o comunitate reproductiv formato comunitate reproductiv format

din populaii care habiteaz un anumit areal idin populaii care habiteaz un anumit areal i

ai cror indivizi se ncrucieaz liber n naturai cror indivizi se ncrucieaz liber n naturcu orice individ de sex opus, dnd natere lacu orice individ de sex opus, dnd natere ladescendeni fertili care menin relativ omogendescendeni fertili care menin relativ omogenfondul de gene al populfondul de gene al populaiei.aiei.


7/22

Spr! /!o"!+$r! /! organ$"-!'Spr! /!o"!+$r! /! organ$"-!'!!"up!r$oar!*"up!r$oar!*bacteriilbacteriile se *nmul$esc vegetativ prine se *nmul$esc vegetativ prindiviiunea celular#diviiunea celular#* $ar "&;$-+u' /! -at!r$a'* $ar "&;$-+u' /! -at!r$a'g!n!t$& !"t! rar >$ nu r!pr!6$nt) un &ara&t!r /!g!n!t$& !"t! rar >$ nu r!pr!6$nt) un &ara&t!r /!

"p!&$!."p!&$!. A&!"t! /$,!r!n5!* 'a &ar! "! a/aug)A&!"t! /$,!r!n5!* 'a &ar! "! a/aug)11::deosebirile 'enotipice mai puin tran3antedeosebirile 'enotipice mai puin tran3anteale bacteriilorale bacteriilor>$>$:: capacitatea mult mai mare de a su'ericapacitatea mult mai mare de a su'erivariaii 'enotipice 3i genotipicevariaii 'enotipice 3i genotipice* "por!"& gra/u'* "por!"& gra/u'/! /$&u'tat! n /!'$-$tar!a "p!&$$'or +a&t!r$!n! >$/! /$&u'tat! n /!'$-$tar!a "p!&$$'or +a&t!r$!n! >$g)"$r!a unu$ pun&t /! 0!/!r! un$tar a"uprag)"$r!a unu$ pun&t /! 0!/!r! un$tar a"upra&on&!ptu'u$ /! "p!&$! +a&t!r$an).&on&!ptu'u$ /! "p!&$! +a&t!r$an).


8/22

SSpeciile bacterienepeciile bacteriene==popu'a5$$ /!popu'a5$$ /!

&!'u'! &u &ara&t!r! ,!not$p$&! &o-un!*&!'u'! &u &ara&t!r! ,!not$p$&! &o-un!*pr$n &ar! /$,!r) "!-n$&at$0 /! a't!pr$n &ar! /$,!r) "!-n$&at$0 /! a't!popu'a5$$.popu'a5$$.

?$!&ar! popu'a5$! &ar! apar5$n! un!$?$!&ar! popu'a5$! &ar! apar5$n! un!$"p!&$$ poart) nu-!'! /!"p!&$$ poart) nu-!'! /! tulpin#tulpin#bacterian#bacterian# ((clon#clon# sausausu3#su3#).).


9/22

$ !"t! g!n!t$& o-og!n).&!'u'! >$ !"t! g!n!t$& o-og!n).


10/22

Subdiviiunile specieiSubdiviiunile specieibacterienebacterieneau aprut ca urmare a sesisriiau aprut ca urmare a sesisrii

unor dieren!e "ntre microorganismele aceleea#i speciiunor dieren!e "ntre microorganismele aceleea#i specii2ubspecia2ubspecia$-p'$&) /$,!r!n5!$-p'$&) /$,!r!n5!ntr!ntr!

&ara&t!r!&ara&t!r!'! &!'! -a$'! &!'! -a$ $-portant!* ,)r)$-portant!* ,)r) &&aaapart!n!n%a 'aapart!n!n%a 'a "p!&$!"p!&$!") ! a't!rat)") ! a't!rat)..

4arianta4arianta((var!var! "au "au tipultipul"! a+at! /! 'a"! a+at! /! 'a,!not$pu',!not$pu'"p!&$!$ pr$ntr:un "$ngur &ara&t!r"p!&$!$ pr$ntr:un "$ngur &ara&t!r/$,!r!n5$a'* "au pr$ntr:un nu-)r r!"tr9n" /!/$,!r!n5$a'* "au pr$ntr:un nu-)r r!"tr9n" /!&ara&t!r!.&ara&t!r!.

n ,un&5$! /! &ara&t!ru' /$,!r!n5$a' "! potn ,un&5$! /! &ara&t!ru' /$,!r!n5$a' "! pot/$"t$ng! -a$ -u't! &at!gor$$ /!/$"t$ng! -a$ -u't! &at!gor$$ /! variantevariante"au"au tipuritipuri ( ta+( ta+!'u'!'u'4.4.


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T$%&'' .T$%&'' .Taxoni inraspecifciTaxoni inraspecifci

*enumirea*enumireataxonuluitaxonului

SinonimSinonim CaracterulCaracteruldieren+ialdieren+ial

S!ro0arS!ro0ar S!rot$pS!rot$p


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Taxonii superiorTaxonii superioriispecieispeciei

+enul+enulr!un!#t!r!un!#t! "p!&$$"p!&$$'!'!nru/$t!nru/$t!* ap)rut! pr$n* ap)rut! pr$n!0o'u%$! /$0!rg!nt) /$ntr:un "tr)-o% &o-un.!0o'u%$! /$0!rg!nt) /$ntr:un "tr)-o% &o-un.Toat!Toat!+a&t!r$$'! >$ au 'o&u' n &a/ru' unu$ g!n* $ar+a&t!r$$'! >$ au 'o&u' n &a/ru' unu$ g!n* $ar$/!nt$&ar!a g!nu'u$ "t) 'a +a6a /$agno"t$&u'u$$/!nt$&ar!a g!nu'u$ "t) 'a +a6a /$agno"t$&u'u$+a&t!r$o'og$& &ur!nt.+a&t!r$o'og$& &ur!nt.

5amilia5amiliagrup!a6) -a$ -u't! g!nur$ nru/$t!.grup!a6) -a$ -u't! g!nur$ nru/$t!.Nu-!roa"! g!nur$ nu au ,o"t n&a/rat! n&) nNu-!roa"! g!nur$ nu au ,o"t n&a/rat! n&) n,a-$'$$.,a-$'$$.

6rdinu6rdinull!"t! un taxon ,or-at /$n ,a-$'$$ nru/$t!!"t! un taxon ,or-at /$n ,a-$'$$ nru/$t!..

ClasaClasa

!"t! un taxon "up!r$or* &ar! grup!a6) -a$!"t! un taxon "up!r$or* &ar! grup!a6) -a$

-u't! or/$n! nru/$t!.-u't! or/$n! nru/$t!. -iviiunea-iviiunea(789lum)(789lum)r!pr!6$nt) o grupar! /!r!pr!6$nt) o grupar! /!

&'a"! nru/$t!.&'a"! nru/$t!. RegnulRegnulgrup!a6) /$0$6$un$'! nru/$t!.grup!a6) /$0$6$un$'! nru/$t!.


13/22

Lu9n/ /r!pt &r$t!r$u /! +a6)Lu9n/ /r!pt &r$t!r$u /! +a6) nivelul denivelul deorganiare biologic#organiare biologic#("u+&!'u'ar* un$&!'u'ar("u+&!'u'ar* un$&!'u'ar

!u&ar$ot "au pro&ar$ot!u&ar$ot "au pro&ar$ot

In 2@B* M u r r a n&a/r!a6)In 2@B* M u r r a n&a/r!a6)+a&t!r$$'! >$ a'g!'! a'+a"tr!:0!r6$+a&t!r$$'! >$ a'g!'! a'+a"tr!:0!r6$(&$ano+a&t!r$$'! n r!gnu'(&$ano+a&t!r$$'! n r!gnu'

7rocar9otae7rocar9otae!!nn 2@@2@@** ; $ t t a ! r &r!!a6) ; $ t t a ! r &r!!a6)

r!gnu'r!gnu' 5ungi5ungi &ar! grup!a6)&ar! grup!a6)

&$up!r&$'! -$&ro"&op$&!.&$up!r&$'! -$&ro"&op$&!. F$ru"ur$'!* au ,o"t n&a/rat! ntr:unF$ru"ur$'!* au ,o"t n&a/rat! ntr:un

r!gn /! "$n! "t)t)tor* r!gnu'r!gn /! "$n! "t)t)tor* r!gnu' 4ira4ira..


14/22

Clasifcarea bacteriilorClasifcarea bacteriilor

U't$-a &'a"$&ar!U't$-a &'a"$&ar! a +a&t!r$$'ora +a&t!r$$'or a ap)rut na ap)rut n 4488n &a/ru' Manua'u'u$ /! S$"t!-at$&) Ha&t!r$an)n &a/ru' Manua'u'u$ /! S$"t!-at$&) Ha&t!r$an)a 'u$ H ! r g ! a 'u$ H ! r g ! #$ &o'a+. (H!rg!" Manua' o,#$ &o'a+. (H!rg!" Manua' o,

S$"t!-at$& Ha&t!r$o'ogS$"t!-at$& Ha&t!r$o'og

(ta+. 7.(ta+. 7.

R!gnu'


15/22

Tabelul Tabelul Clasifcarea bacteriilor la nivelul taxonilorClasifcarea bacteriilor la nivelul taxonilor

superiorisuperiori

&/' 01C$21T$&&/' 01C$21T$&*iviiunea (0h3lum)*iviiunea (0h3lum) ClasaClasa

4.4. racilicutesracilicutesK +a&t!r$$ Gra-K +a&t!r$$ Gra-n!gat$0!n!gat$0!

II..2.2. !cotobacteria!cotobacteriaI.4.I.4."nox#photobacteria"nox#photobacteria

I.7.I.7. $x#photobacteria$x#photobacteria44.44. %irmicutes%irmicutesK +a&t!r$$ Gra-K +a&t!r$$ Gra-po6$t$0!po6$t$0!

II.2.II.2. %irmibacteria%irmibacteria

II.4.II.4. &hallobacteria&hallobacteria444.444. &enericutes&enericutesK +a&t!r$$K +a&t!r$$'$p"$t! /! p!r!t! &!'u'ar'$p"$t! /! p!r!t! &!'u'ar

III.2.III.2. 'ollicutes'ollicutes

45.45. 'endosicutes'endosicutesK p!r!t!K p!r!t!&!'u'ar '$p"$t /! "tratu' /!&!'u'ar '$p"$t /! "tratu' /!p!pt$/og'$&anp!pt$/og'$&an

IF.2.IF.2."rchaeobacteria"rchaeobacteria


16/22

/16&/C'$T$/16&/C'$T$%$CT&44'1%$CT&44'1

(numirea #tiin!ifc)(numirea #tiin!ifc)Con,or- nor-!'orCon,or- nor-!'or $nt!rna%$ona'!$nt!rna%$ona'!/! no-!n&'atur) n +$o'og$!*/! no-!n&'atur) n +$o'og$!*

"p!&$a"p!&$a!"t! nu-$t)!"t! nu-$t)pr$n /ou)pr$n /ou)&u0$nt! 'at$n$6at! &ar!&u0$nt! 'at$n$6at! &ar!&ara&t!r$6!a6)&ara&t!r$6!a6) &9t -a$&9t -a$ "$nt!t$&"$nt!t$&

-$&roorgan$"-u' r!"p!&t$0-$&roorgan$"-u' r!"p!&t$0..EEx!-p'x!-p'!1!1 !treptococcus!treptococcusp#ogenes, !taph#lococcus aureusp#ogenes, !taph#lococcus aureus


17/22

$ !"t!>$ !"t!$n"p$rat /!$n"p$rat /!11

caractere mor'ologicecaractere mor'ologice* ,r!&0!nt ,or-a >$ -o/u'* ,r!&0!nt ,or-a >$ -o/u'/! grupar! (/! grupar! ((acillus(acillus = +a"ton -$&3= +a"ton -$&3!taph#lococcus!taph#lococcus= &$or&;$n! /! "trugur!3= &$or&;$n! /! "trugur!3 !arcina!arcina= pa&;!t !t&.* a"o&$at! un!or$ &u ;a+$tatu'= pa&;!t !t&.* a"o&$at! un!or$ &u ;a+$tatu'natura' a' +a&t!r$!$ (natura' a' +a&t!r$!$ ()actobacillus)actobacillus= +a"tona> /$n= +a"tona> /$n'apt!3'apt!3

numele bacteriologuluinumele bacteriologului&ar! a $6o'at pr$-u'&ar! a $6o'at pr$-u'+a&t!r$a r!"p!&t$0) "au a a0ut -!r$t! /!o"!+$t! n+a&t!r$a r!"p!&t$0) "au a a0ut -!r$t! /!o"!+$t! n"tu/$u' !$1"tu/$u' !$1 (rucella(rucella(Hru&!*(Hru&!* *asteurella*asteurella(


18/22

C!' /!:a' /o$'!a &u09nt /!nu-!>t!C!' /!:a' /o$'!a &u09nt /!nu-!>t!speciaspecia!!E'E' "!"! poat!poat! r!,!r$ 'ar!,!r$ 'a11

un caracterun caracter-or,o'og$&* &u'tura' "au +$o&;$-$&-or,o'og$&* &u'tura' "au +$o&;$-$&((!taph#lococcus aureus!taph#lococcus aureus** )actobacillus)actobacillusacidophilusacidophilus!t&.3!t&.3

gada receptiv#gada receptiv#'a &ar! pro/u&! $n,!&5$$'a &ar! pro/u&! $n,!&5$$

(((rucella suis(rucella suis** !treptococcus eui!treptococcus eui** '#coplasma'#coplasmagallinarumgallinarum!t&.3!t&.3

boalaboala* un* un simptomsimptom"au o"au o leiuneleiunecaracteristic#caracteristic#((-lostridium tetani-lostridium tetani** '#coplasma'#coplasma

agalactiaeagalactiae** )isteria monoc#togenes)isteria monoc#togenes!t&.3!t&.3numele bacteriologuluinumele bacteriologului&ar! a /!"&r$" pr$-u'&ar! a /!"&r$" pr$-u'+a&t!r$a (+a&t!r$a ("ctinobacillus lignieresii"ctinobacillus lignieresii** -lostridium-lostridiumchauvoeichauvoei** -oxiella burneti-oxiella burneti!t&..!t&..


19/22

Nu-!roa"! "p!&$$ +a&t!r$!n! au >$Nu-!roa"! "p!&$$ +a&t!r$!n! au >$nu-! &o-un!* ,o'o"$t! /!"!or$ nnu-! &o-un!* ,o'o"$t! /!"!or$ n0or+$r!a &ur!nt)* &a /! !x!-p'u 10or+$r!a &ur!nt)* &a /! !x!-p'u 1+a&$'u' p$o&$an$& "au +a&$'u' puro$u'u$+a&$'u' p$o&$an$& "au +a&$'u' puro$u'u$a'+a"tru* p!ntrua'+a"tru* p!ntru 7seudomonas7seudomonasaeruginosaaeruginosa3 +a&$'u' 'u$ o&; "au3 +a&$'u' 'u$ o&; "au

H..* p!ntruH..* p!ntru :9cobacterium:9cobacteriumtuberculosistuberculosis3 +a&$'u' t!tano"u'u$*3 +a&$'u' t!tano"u'u$*p!ntrup!ntru Clostridium tetaniClostridium tetani!t&.!t&.


20/22

66tulpin# bacterian#tulpin# bacterian#&ar! nu a&ar! nu a,o"t n&) $/!nt$&at),o"t n&) $/!nt$&at) !"t!!"t!nu-$nu-$t)t)iolat;iolat;$ar /up) n&a/rar!a n g!n*$ar /up) n&a/rar!a n g!n*p9n) 'a pr!&$6ar!a "p!&$!$*p9n) 'a pr!&$6ar!a "p!&$!$* poart)poart)/!nu-$r!a g!nu'u$ ur-at) /!/!nu-$r!a g!nu'u$ ur-at) /!a+r!0$!r!a t!r-!nu'u$a+r!0$!r!a t!r-!nu'u$ species< sp!species< sp!

p!ntru "$ngu'ar >$p!ntru "$ngu'ar >$ spp!spp!p!ntru p'ura'p!ntru p'ura'(!x.(!x. 2treptococcus sp! 2almonella2treptococcus sp! 2almonellaspp!)!spp!)!


21/22

2ubspeciile 3i variantele2ubspeciile 3i variantele "! /!nu-!"&"! /!nu-!"&a/)ug9n/ 'a nu-!'! "p!&$!$* t!r-!n$$a/)ug9n/ 'a nu-!'! "p!&$!$* t!r-!n$$subspeciessubspecies(pr!"&urtat(pr!"&urtat subspsubsp.),.),r!"p!&t$0r!"p!&t$0 varietasvarietas(pr!"&urtat(pr!"&urtat varvar.),.),ur-a5$ /! nu-!'! "u+"p!&$!$ "au 0ar$ant!$ur-a5$ /! nu-!'! "u+"p!&$!$ "au 0ar$ant!$r!"p!&t$0!1r!"p!&t$0!1 Camp9lobacter 'etus subsp!Camp9lobacter 'etus subsp!'etus'etus33

Far$ant!'! "! -a$ not!a6) &uFar$ant!'! "! -a$ not!a6) &u ci'reci're('a('a!treptococcus pneumoniae!treptococcus pneumoniae>$>$ (rucella spp(rucella spp.*.*literelitere('a('a -lostridium perfringens-lostridium perfringens>$>$ -lostridium-lostridiumbotulinumbotulinum* "au pr$n "p!&$&ar!a* "au pr$n "p!&$&ar!a antigenilor 6antigenilor 63i =3i =* ('a* ('a +scherichia coli+scherichia coli>$>$ !almonella spp!almonella spp....


22/22

6rdinul6rdinul>$>$ 'amilia'amilia "unt /!nu-$t! &u"unt /!nu-$t! &unu-!'! g!nu'u$ &!'u$ -a$ r!pr!6!ntat$0 'anu-!'! g!nu'u$ &!'u$ -a$ r!pr!6!ntat$0 'a&ar! "! a/aug) "ux!'! **&ar! "! a/aug) "ux!'! **ales;ales;p!ntru or/$np!ntru or/$n>$>$ ,,,,aceaeaceae77p!ntru ,a-$'$! (!x. g!nu'p!ntru ,a-$'$! (!x. g!nu'

r!pr!6!ntat$0r!pr!6!ntat$0 ic/ettsiaic/ettsia* or/$nu'* or/$nu'ic/ettsialesic/ettsiales* ,a-$'$a* ,a-$'$a ic/ettsiaceae0ic/ettsiaceae0.. >ncreng#tura>ncreng#tura(diviiunea)(diviiunea)>$>$ clasaclasaK taxon$K taxon$

/! rang "up!r$or K au /!nu-$r$ &on"t$tu$t!/! rang "up!r$or K au /!nu-$r$ &on"t$tu$t!/$ntr:un nu-! g!n!r$& >$ /$0!r"!/$ntr:un nu-! g!n!r$& >$ /$0!r"! sufxesufxe. La. La

ora a&tua')* n taxono-$a +a&t!r$an) "!ora a&tua')* n taxono-$a +a&t!r$an) "!-an$,!"t) t!n/$n5a /! a "! op!ra /$n &! n &!-an$,!"t) t!n/$n5a /! a "! op!ra /$n &! n &!-a$ pu5$n &u a&!"t! &at!gor$$ taxono-$&!*-a$ pu5$n &u a&!"t! &at!gor$$ taxono-$&!*/$n &au6a r!'at$0$t)5$$ >$ $n!&$!n5!$ 'or./$n &au6a r!'at$0$t)5$$ >$ $n!&$!n5!$ 'or.

curs pp4 sistem

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